*{margin:0;padding:0;box-sizing:border-box}html{font-size:16px;line-height:1.6;scroll-behavior:smooth}body{font-family:inter,-apple-system,BlinkMacSystemFont,segoe ui,Roboto,sans-serif;color:#1f2937;background-color:#fff;line-height:1.6}.skip-link{position:absolute;top:-40px;left:6px;background:#2563eb;color:#fff;padding:8px;text-decoration:none;border-radius:4px;z-index:9999}.skip-link:focus{top:6px}.container{max-width:1200px;margin:0 auto;padding:0 1rem}.site-header{background:#fff;border-bottom:1px solid #e5e7eb;position:sticky;top:0;z-index:100;box-shadow:0 1px 3px rgba(0,0,0,.1)}.navbar{padding:1rem 0}.navbar .container{display:flex;justify-content:space-between;align-items:center}.navbar-brand{display:flex;align-items:center}.navbar-brand a{text-decoration:none;color:#1f2937}.site-title{font-size:1.5rem;font-weight:700;color:#2563eb}.logo{height:60px;width:auto}.navbar-menu{display:flex;align-items:center;gap:2rem}.navbar-item{text-decoration:none;color:#4b5563;font-weight:500;padding:.5rem 1rem;border-radius:6px;transition:all .2s ease}.navbar-item:hover,.navbar-item.is-active{background-color:#f3f4f6;color:#2563eb}.navbar-burger{display:none;flex-direction:column;width:24px;height:24px;background:0 0;border:none;cursor:pointer}.navbar-burger span{display:block;height:2px;width:100%;background-color:#4b5563;margin:3px 0;transition:.3s}.search-toggle{background:0 0;border:none;cursor:pointer;color:#4b5563;padding:.5rem;border-radius:6px;transition:all .2s ease}.search-toggle:hover{background-color:#f3f4f6;color:#2563eb}.search-overlay{position:fixed;top:0;left:0;right:0;bottom:0;background:rgba(0,0,0,.8);z-index:1000;display:none;padding:2rem}.search-overlay.active{display:flex;flex-direction:column;justify-content:flex-start;align-items:center;padding-top:10%}.search-container{position:relative;width:100%;max-width:600px;margin-bottom:2rem}#search-input{width:100%;padding:1rem 1.5rem;font-size:1.125rem;border:none;border-radius:12px;background:#fff;box-shadow:0 4px 6px rgba(0,0,0,.1)}#search-close{position:absolute;top:50%;right:1rem;transform:translateY(-50%);background:0 0;border:none;cursor:pointer;color:#6b7280}.search-results{width:100%;max-width:600px;background:#fff;border-radius:12px;max-height:400px;overflow-y:auto;box-shadow:0 4px 6px rgba(0,0,0,.1)}.hero{background:linear-gradient(135deg,#667eea 0%,#764ba2 100%);color:#fff;padding:6rem 0;text-align:center}.hero-title{font-size:3.5rem;font-weight:700;margin-bottom:1.5rem;line-height:1.2}.hero-subtitle{font-size:1.5rem;margin-bottom:3rem;opacity:.9;max-width:600px;margin-left:auto;margin-right:auto}.hero-stats{display:flex;justify-content:center;gap:4rem;margin-top:3rem}.stat{text-align:center;padding:1rem}.stat-number{display:block;font-size:3rem;font-weight:700;color:#fbbf24;margin-bottom:.5rem}.stat-label{font-size:1rem;opacity:.9;font-weight:500}.featured-article{padding:6rem 0;background:#f9fafb}.section-title{font-size:2.5rem;font-weight:700;margin-bottom:3rem;text-align:center;color:#1f2937}.featured-post{display:grid;grid-template-columns:1fr 1fr;gap:4rem;background:#fff;border-radius:16px;overflow:hidden;box-shadow:0 8px 25px rgba(0,0,0,.1);margin-bottom:2rem}.featured-image{height:500px;overflow:hidden;display:flex;align-items:center;justify-content:center;background:#f9fafb}.featured-image img{width:100%;height:100%;object-fit:cover}.featured-content{padding:3rem}.post-meta{display:flex;gap:1rem;margin-bottom:1rem;font-size:.875rem;color:#6b7280}.category a{background:#dbeafe;color:#1d4ed8;padding:.25rem .75rem;border-radius:999px;text-decoration:none;font-weight:500}.research-type{background:#f3e8ff;color:#7c3aed;padding:.25rem .75rem;border-radius:999px;font-weight:500}.post-title{font-size:2rem;font-weight:700;margin-bottom:1.5rem;line-height:1.3}.post-title a{text-decoration:none;color:#1f2937}.post-title a:hover{color:#2563eb}.post-summary{color:#4b5563;margin-bottom:2rem;line-height:1.7;font-size:1.125rem}.key-findings{background:#f0f9ff;padding:2rem;border-radius:12px;margin-bottom:2rem;border-left:6px solid #0ea5e9}.key-findings h4{color:#0369a1;margin-bottom:.5rem;font-weight:600}.key-findings ul{list-style:none;padding-left:0}.key-findings li{position:relative;padding-left:1.5rem;margin-bottom:.5rem;color:#1e40af}.key-findings li:before{content:"•";color:#0ea5e9;position:absolute;left:0;font-weight:700}.post-footer{display:flex;justify-content:space-between;align-items:flex-start;gap:2rem;flex-wrap:wrap}.read-more{background:#2563eb;color:#fff;padding:1rem 2rem;text-decoration:none;border-radius:8px;font-weight:500;transition:background .2s ease;white-space:nowrap;flex-shrink:0}.read-more:hover{background:#1d4ed8}.species-tags{display:flex;gap:.5rem;flex-wrap:wrap;align-items:flex-start}.species-tag{background:#ecfdf5;color:#065f46;padding:.25rem .5rem;border-radius:999px;font-size:.75rem;font-weight:500}.recent-articles{padding:6rem 0}.articles-grid{display:grid;grid-template-columns:repeat(auto-fit,minmax(350px,1fr));gap:2.5rem;margin-bottom:4rem}.article-card{background:#fff;border-radius:16px;overflow:hidden;box-shadow:0 4px 15px rgba(0,0,0,.1);transition:transform .3s ease,box-shadow .3s ease}.article-card:hover{transform:translateY(-4px);box-shadow:0 8px 30px rgba(0,0,0,.15)}.article-image img{width:100%;height:200px;object-fit:cover}.article-content{padding:2rem}.article-meta{display:flex;gap:1rem;margin-bottom:1rem;font-size:.875rem;color:#6b7280;flex-wrap:wrap}.article-title{font-size:1.25rem;font-weight:700;margin-bottom:1rem;line-height:1.4}.article-title a{text-decoration:none;color:#1f2937}.article-title a:hover{color:#2563eb}.article-summary{color:#4b5563;margin-bottom:1.5rem;line-height:1.6;font-size:1rem}.article-footer{display:flex;justify-content:space-between;align-items:center;font-size:.875rem;color:#6b7280}.read-time{font-weight:500}.view-all{text-align:center}.btn{display:inline-block;padding:.75rem 1.5rem;text-decoration:none;border-radius:6px;font-weight:500;transition:all .2s ease;border:none;cursor:pointer}.btn-primary{background:#2563eb;color:#fff}.btn-primary:hover{background:#1d4ed8;transform:translateY(-1px)}.categories-overview{padding:6rem 0;background:#f9fafb}.categories-grid{display:grid;grid-template-columns:repeat(auto-fit,minmax(280px,1fr));gap:2rem}.category-card{background:#fff;padding:2.5rem 2rem;border-radius:16px;text-align:center;box-shadow:0 4px 15px rgba(0,0,0,.1);transition:transform .3s ease,box-shadow .3s ease}.category-card:hover{transform:translateY(-4px);box-shadow:0 8px 25px rgba(0,0,0,.15)}.category-card h3{margin-bottom:1rem}.category-card h3 a{text-decoration:none;color:#1f2937;font-weight:700;font-size:1.25rem}.category-card h3 a:hover{color:#2563eb}.category-count{color:#6b7280;font-size:1rem;margin-bottom:1rem;font-weight:600}.category-description{color:#4b5563;font-size:1rem;line-height:1.6}.newsletter{background:#1f2937;color:#fff;padding:6rem 0;text-align:center}.newsletter-content h2{font-size:2.5rem;margin-bottom:1.5rem;font-weight:700}.newsletter-content p{font-size:1.25rem;margin-bottom:3rem;opacity:.9;max-width:500px;margin-left:auto;margin-right:auto}.newsletter-form{display:flex;justify-content:center;gap:1rem;max-width:400px;margin:0 auto}.newsletter-form input{flex:1;padding:.75rem 1rem;border:none;border-radius:6px;font-size:1rem}.newsletter-form button{padding:.75rem 1.5rem;background:#2563eb;color:#fff;border:none;border-radius:6px;font-weight:500;cursor:pointer;transition:background .2s ease}.newsletter-form button:hover{background:#1d4ed8}.single-post{padding:2rem 0}.article-header{margin-bottom:2rem}.article-title{font-size:2.5rem;font-weight:700;line-height:1.2;margin-bottom:1rem;color:#1f2937}.article-summary{font-size:1.125rem;color:#4b5563;margin-bottom:1.5rem;line-height:1.6}.research-details{display:grid;grid-template-columns:repeat(auto-fit,minmax(200px,1fr));gap:1rem;margin-bottom:1.5rem;padding:1.5rem;background:#f9fafb;border-radius:8px}.detail-item{display:flex;flex-direction:column}.detail-label{font-weight:600;color:#374151;font-size:.875rem}.detail-value{color:#1f2937;font-weight:500}.article-taxonomies{margin-bottom:1.5rem}.taxonomy-group{display:flex;align-items:center;gap:1rem;margin-bottom:1rem}.taxonomy-label{font-weight:600;color:#374151;min-width:80px}.taxonomy-items{display:flex;gap:.5rem;flex-wrap:wrap}.tag{background:#f3f4f6;color:#374151;padding:.25rem .75rem;border-radius:999px;text-decoration:none;font-size:.875rem;font-weight:500;transition:all .2s ease}.tag:hover{background:#e5e7eb}.featured-image{margin-bottom:2rem}.featured-image img{width:100%;height:400px;object-fit:contain;border-radius:12px;background:#f9fafb}.key-findings-box{background:#f0f9ff;padding:2rem;border-radius:12px;margin-bottom:2rem;border-left:4px solid #0ea5e9}.key-findings-box h3{color:#0369a1;margin-bottom:1rem;font-weight:600}.key-findings-box ul{list-style:none;padding-left:0}.key-findings-box li{position:relative;padding-left:1.5rem;margin-bottom:.75rem;color:#1e40af;line-height:1.6}.key-findings-box li:before{content:"✓";color:#0ea5e9;position:absolute;left:0;font-weight:700}.article-content{font-size:1.125rem;line-height:1.8;color:#374151;margin-bottom:3rem}.article-content h2{font-size:1.5rem;font-weight:600;margin:2rem 0 1rem;color:#1f2937}.article-content h3{font-size:1.25rem;font-weight:600;margin:1.5rem 0 1rem;color:#1f2937}.article-content p{margin-bottom:1.5rem}.article-content ul,.article-content ol{margin-bottom:1.5rem;padding-left:1.5rem}.article-content li{margin-bottom:.5rem}.article-content blockquote{border-left:4px solid #e5e7eb;padding-left:1.5rem;margin:1.5rem 0;font-style:italic;color:#6b7280}.research-citation{background:#f9fafb;padding:2rem;border-radius:12px;margin-bottom:2rem;border:1px solid #e5e7eb}.research-citation h3{color:#1f2937;margin-bottom:1rem;font-weight:600}.citation-text{font-style:italic;color:#4b5563;margin-bottom:1rem;line-height:1.6}.doi-link a{color:#2563eb;text-decoration:none;font-weight:500}.doi-link a:hover{text-decoration:underline}.social-sharing{margin-bottom:3rem}.social-sharing h3{margin-bottom:1rem;color:#1f2937}.sharing-buttons{display:flex;gap:1rem;flex-wrap:wrap}.share-btn{display:flex;align-items:center;gap:.5rem;padding:.75rem 1rem;text-decoration:none;border-radius:6px;font-weight:500;transition:all .2s ease;border:none;cursor:pointer}.share-btn.twitter{background:#1da1f2;color:#fff}.share-btn.facebook{background:#1877f2;color:#fff}.share-btn.linkedin{background:#0a66c2;color:#fff}.share-btn.copy-link{background:#6b7280;color:#fff}.share-btn:hover{transform:translateY(-1px);box-shadow:0 2px 4px rgba(0,0,0,.1)}.related-articles{margin-bottom:3rem}.related-articles h3{margin-bottom:1.5rem;color:#1f2937}.related-grid{display:grid;grid-template-columns:repeat(auto-fit,minmax(250px,1fr));gap:1.5rem}.related-article{background:#fff;border-radius:8px;overflow:hidden;box-shadow:0 2px 4px rgba(0,0,0,.1);transition:transform .2s ease}.related-article:hover{transform:translateY(-2px)}.related-image img{width:100%;height:150px;object-fit:cover}.related-content{padding:1rem}.related-meta{display:flex;gap:1rem;margin-bottom:.5rem;font-size:.875rem;color:#6b7280}.related-content h4{margin-bottom:.5rem;font-size:1rem}.related-content h4 a{text-decoration:none;color:#1f2937}.related-content h4 a:hover{color:#2563eb}.related-content p{color:#4b5563;font-size:.875rem;line-height:1.5}.site-footer{background:#1f2937;color:#fff;padding:3rem 0 1rem}.footer-content{display:grid;grid-template-columns:repeat(auto-fit,minmax(250px,1fr));gap:2rem;margin-bottom:2rem}.footer-section h3{margin-bottom:1rem;color:#f9fafb}.footer-section h4{margin-bottom:1rem;color:#d1d5db}.footer-section p{color:#9ca3af;line-height:1.6;margin-bottom:1rem}.footer-section ul{list-style:none}.footer-section li{margin-bottom:.5rem}.footer-section a{color:#9ca3af;text-decoration:none;transition:color .2s ease}.footer-section a:hover{color:#f9fafb}.social-links{display:flex;gap:1rem;margin-top:1rem}.social-links a{color:#9ca3af;transition:color .2s ease}.social-links a:hover{color:#2563eb}.footer-bottom{display:flex;justify-content:space-between;align-items:center;padding-top:2rem;border-top:1px solid #374151;color:#9ca3af;font-size:.875rem}.footer-links{display:flex;gap:1rem}.footer-links a{color:#9ca3af;text-decoration:none;transition:color .2s ease}.footer-links a:hover{color:#f9fafb}.list-page{padding:2rem 0}.page-header{text-align:center;margin-bottom:3rem}.page-title{font-size:2.5rem;font-weight:700;margin-bottom:1rem;color:#1f2937}.page-description{font-size:1.125rem;color:#4b5563;margin-bottom:1rem}.page-stats{display:flex;justify-content:center;gap:2rem;color:#6b7280;font-size:.875rem}.filters{display:flex;gap:1rem;margin-bottom:2rem;flex-wrap:wrap}.filter-group{display:flex;align-items:center;gap:.5rem}.filter-group label{font-weight:500;color:#374151}.filter-group select{padding:.5rem 1rem;border:1px solid #d1d5db;border-radius:6px;background:#fff;color:#374151}.key-findings-preview{background:#f0f9ff;padding:1rem;border-radius:6px;margin-bottom:1rem;border-left:3px solid #0ea5e9;font-size:.875rem}.key-findings-preview strong{color:#0369a1}.article-stats{display:flex;gap:1rem;align-items:center;font-size:.875rem;color:#6b7280}.journal{font-style:italic}.species-tag.more{background:#f3f4f6;color:#6b7280}.no-articles{text-align:center;padding:4rem 0;color:#6b7280}.no-articles h3{font-size:1.5rem;margin-bottom:1rem;color:#374151}.no-articles p{margin-bottom:2rem}.pagination{margin-top:3rem;text-align:center}.pagination-info{margin-bottom:1rem;color:#6b7280;font-size:.875rem}.pagination-nav{display:flex;justify-content:center;gap:.5rem;flex-wrap:wrap}.pagination-btn{padding:.5rem 1rem;text-decoration:none;border-radius:6px;color:#4b5563;background:#fff;border:1px solid #d1d5db;transition:all .2s ease}.pagination-btn:hover{background:#f9fafb;border-color:#9ca3af}.pagination-btn.current{background:#2563eb;color:#fff;border-color:#2563eb}.sidebar{background:#f9fafb;padding:2rem;border-radius:12px;margin-top:2rem}.sidebar-section{margin-bottom:2rem}.sidebar-section h3{margin-bottom:1rem;color:#1f2937;font-size:1.125rem}.sidebar-section ul{list-style:none}.sidebar-section li{display:flex;justify-content:space-between;align-items:center;margin-bottom:.5rem}.sidebar-section a{color:#4b5563;text-decoration:none;transition:color .2s ease}.sidebar-section a:hover{color:#2563eb}.count{color:#9ca3af;font-size:.875rem}@media(max-width:768px){.hero-title{font-size:2rem}.hero-stats{flex-direction:column;gap:1rem}.featured-post{grid-template-columns:1fr}.articles-grid{grid-template-columns:1fr}.categories-grid{grid-template-columns:1fr}.newsletter-form{flex-direction:column}.article-title{font-size:2rem}.research-details{grid-template-columns:1fr}.sharing-buttons{flex-direction:column}.footer-bottom{flex-direction:column;gap:1rem;text-align:center}.navbar-menu{display:none;position:absolute;top:100%;left:0;right:0;background:#fff;flex-direction:column;padding:1rem;box-shadow:0 2px 4px rgba(0,0,0,.1)}.navbar-menu.active{display:flex}.navbar-burger{display:flex}.filters{flex-direction:column}.pagination-nav{flex-wrap:wrap}.page-stats{flex-direction:column;gap:.5rem}}@media(max-width:480px){.container{padding:0 .5rem}.hero-title{font-size:1.5rem}.hero-subtitle{font-size:1rem}.article-title{font-size:1.5rem}.featured-content{padding:1rem}.article-content{font-size:1rem}.related-grid{grid-template-columns:1fr}}@media print{.site-header,.site-footer,.social-sharing,.related-articles,.newsletter{display:none}.single-post{padding:0}.article-content{font-size:12pt;line-height:1.5}.article-title{font-size:18pt}.key-findings-box{background:#f5f5f5;border:1px solid #ccc}}